Professor Susan E. Evans


The group has a primary research interest in herpetology, with the major focus on the evolution, systematics and palaeobiogeography of small reptiles and amphibians. Our work incorporates both fossil and living genera.

Herpetology is the study of reptiles and amphibians, living and extinct. Our group has particular interests in the evolution, diversification, functional morphology and relationships of lizards and their relatives; in the early evolution of living amphibian groups; and in a small range of exclusively fossil clades. Although our principal focus is palaeontological, building hypotheses on the basis of fossil material requires an understanding of modern musculoskeletal structure, function and ontogeny, as well as phylogenetic relationships. We use a mixture of techniques including cladistics, morphometrics, scanning electron microscopy, and anything else that sheds light on the major questions at hand – namely when, where and how these groups diversified, how modern characteristics evolved and in what order, how the groups diversified through changing continental configurations, and why some groups became extinct while others survived and flourished. The canvas on which we work is necessarily a broad one, but each individual project aims to fill in one further piece of the fine detail.

Lizards and lissamphibians generally have small, delicately built skeletons and are not ideal candidates for fossilisation. Consequently, their fossil remains are rare, and localities with suitable preservational conditions are scattered around the world. In addition to several UK based projects, we are currently collaborating with colleagues in Continental Europe (Spain, France, Poland), Japan, India, Australia and North America, involving fossil deposits ranging in age from Early Triassic (circa 240 Myrs BP) to Recent (subfossil).

However, fossils cannot be studied in isolation, particularly if they have close living relatives. Their incorporation into phylogenetic trees based on modern taxa is important in the testing of the phylogenetic hypotheses represented by those trees. Moreover, many fossil taxa are represented by single specimens, frequently incomplete, and it is important to know how representative such specimens are of the taxon as a whole, i.e. how much skeletal variation (ontogenetic, sexual, regional) occurs within modern species and how do we distinguish such intraspecific variation from phylogenetically significant variation characterising discrete taxa. Such studies have been made for a few vertebrate groups (e.g. primates), but very little has been done on reptiles and amphibians. There are also many questions relating to the individual characters used in phylogenetic analysis, particularly with respect to the influence of function and development on skeletal features, and also the independence or interdependence of apparently discrete features.

Albanerpetontid amphibian, 
U/V light showing soft tissues. 

Las Hoyas, Spain.

 


Field crew at Shokawa site, Gifu Prefecture, Japan.


 

Sample publications:

McGOWAN, G. and EVANS, S.E. 1995. Albanerpetontid amphibians from the early Cretaceous of Spain. Nature, 373:143-145.
EVANS, S.E. & SIGOGNEAU-RUSSELL, D. 1997. New sphenodontians (Diapsida: Lepidosauria: Rhynchocephalia) from the Early Cretaceous of North Africa. Journal of Vertebrate Palaeontology, 17: 45-51.
EVANS, S.E. and CHURE, D. 1998. Paramacellodid lizard skulls from the Jurassic Morrison Formation at Dinosaur National Monument, Utah. Journal of Vertebrate Paleontology, 18: 99-114.
EVANS, S.E. & BARBADILLO, L.J. 1998. An unusual lizard (Reptilia, Squamata) from the Early Cretaceous of Las Hoyas, Spain. Zoological Journal of the Linnean Society, 124: 235-266.
EVANS, S.E. & BORSUK-BIALYNICKA, M. 1998. A stem-frog from the Early Triassic of Poland. Acta Paleontologia Polonica, 43, 4: 573-580.
EVANS SE & MANABE M. 1999. Early Cretaceous lizards from the Okurodani Formation of Japan. Geobios, 32: 889-899.
BARAHONA, F., EVANS, S.E., MATEO, LOPEZ-JURADO, L-P. 2000. Endemism, gigantism and extinction in island lizards: the genus Gallotia on the Canary Islands. Journal of Zoology, 250: 373-388.
MILNER, A.C., MILNER, A.R. and EVANS, S.E. 2000. Global changes and biota: amphibians, reptiles and birds. In, Culver, S. and Rawson, P. (Eds), Biotic response to global change: the last 145 million years, Chapman and Hall, Publishers.
GAO, K., EVANS, S.E., NORRELL, M. and JI, S. 2000. Exceptional material of a semi-aquatic reptile from the Cretaceous of China: the resolution of an enigma. Journal of Vertebrate Paleontology, 20: 417-421.
EVANS, S.E. and SIGOGNEAU-RUSSELL, D. 2001. A stem-group caecilian (Lissamphibia: Gymnophiona) from the Lower Cretaceous of North Africa.  Palaeontology, 44 (2): 259-273.
EVANS, S.E. PRASAD, G.V.R. & MANHAS, B. 2002. An acrodont iguanian from the Mesozoic Kota Formation of  India.  Journal of Vertebrate Paleontology, 22: 299-312.
EVANS, S.E. & SEARLE, B. 2002. The lepidosaurian assemblage of the Purbeck Limestone Group. Special Papers in Palaeontology, 68: 145-159.
EVANS, S.E. & McGOWAN, G. 2002.  An amphibian assemblage from the Purbeck Limestone Group. Special Papers in Palaeontology, 68: 103-119.
JONES, M., EVANS, S.E. & SIGOGNEAU-RUSSELL, D. 2003. Fossil frogs from the Early Cretaceous of Morocco. Annals of the Carnegie Museum, 72: 65-97.
GARDNER, J., EVANS, S.E. & SIGOGNEAU-RUSSELL, D. 2003. Albanerpetontid amphibians from the Early Cretaceous of North Africa.  Acta Palaeontologia Polonica, 48: 301-319.
BELL, C.J., EVANS, S.E. & MAISANO, J. 2003. On the skull of the gymnophthalmid lizard (Squamata: Gymnophthalmidae) Neusticurus ecpleopus. Zoological Journal of the Linnean Society, 139: 283-304.
EVANS, S.E.  2003. At the feet of the dinosaurs: the origin, evolution and early diversification of squamate reptiles (Lepidosauria: Diapsida). Biological Reviews, Cambridge, 78 (4): 513-551.
KRAUSE, D., EVANS, S.E. & GAO, K.  2003. First definitive record of a Mesozoic lizard from Madagascar. Journal of Vertebrate Paleontology, 23 (4): 842-856.
BORSUK-BIALYNICKA, M. & EVANS, S.E. 2003. A basal archosauriform from the Early Triassic of Poland. Acta Palaeontologica Polonica, 48 (4): 649-652.
EVANS, S.E., BARRETT, P. and WARD, D. 2004. First evidence of lizards and amphibians from the Wessex Formation of the Isle of Wight. Proceedings of the Geological Association, 115: 1-9.
EVANS, S.E., RAIA, P. and BARBERA, C. 2004. New lizards and sphenodontians from the Early Cretaceous of Italy. Acta Palaeontologica Polonica, 49 (3): 393-408.
MÜLLER, J., RENESTO, S. & EVANS, S.E. 2005.  The Triassic marine reptile Endennasaurus from Italy (Diapsida: Thalattosauriformes). Palaeontology, 48: 15-30.
EVANS, S.E. & KLEMBARA, J. 2005. A choristoderan reptile (Reptilia; Diapsida) from the Lower Miocene of northwest Bohemia (Czech Republic). J. Vertebrate Paleontology, 25: 168-181.
EVANS, S.E., LALLY, C. , CHURE, D. , ELDER, A. & MAISANO, J. 2005. A new fully metamorphosed salamander from the Late Jurassic of North America. Zoological J. Linnean Society, 143: 599-616.
EVANS, S.E. and WANG, Y. 2005. Dalinghosaurus, a lizard from the Early Cretaceous Jehol Biota of northeast China. Acta Palaeontologica Polonica, 50 (in press).
EVANS, S.E., WANG, Y. & LI, C. 2005. The Early Cretaceous Chinese lizard, Yabeinosaurus: resolving an enigma. Journal of Systematic Palaeontology, 3 (4): (in press)
WANG, Y., JONES, M.E.H. & EVANS, S.E. 2006. A juvenile anuran from the Lower Cretaceous Jiufotang Formation, Liaoning, China. Cretaceous Research. (in press) APPENDIX: Frog hind limb proportions